Mother Nature struggles to maintain equilibrium in a changing world while fire, disease, logging, human displacement, and war repeatedly destroy forests of centuries-old trees. Split Toe, a deer chosen at birth for a unique education, travels through time to understand the interconnected workings of a Michigan forest. He meets humans along the way: Ice Age hunters who trap and kill a mastodon; Mukwoh, a young Ojibwe hunter who stalks Split Toe through swamp and forest; loggers clear-cutting Michigan's white pines; Edra, a woman advocating for the trees; Angus and Grace, pioneers who become a first generation of family farmers; scientists from the future studying the impact of nuclear radiation.
Split Toe witnesses two hundred years of conflict building between modern humans-who fight to control the natural world-and Mother Nature, who repeatedly reaches for balance. He wonders whether human ways will ultimately overpower Mother Nature, until he meets a boy who changes everything.